Fireweed - Epilobium angustifolium L. - Cyprus

 invasive species

Chamerion angustifolium, commonly known as fireweed (mainly in North America), great willow-herb (some parts of Canada), orrosebay willowherb (mainly in Britain), is a perennial herbaceous plant in the willowherb family Onagraceae.

It is native throughout the temperate Northern Hemisphere, including large parts of the boreal forests.

This species has been placed in the genus Chamerion (sometimes, incorrectly, given as Chamaenerion) rather than Epilobium based on several morphological distinctions: spiral (rather than opposite or whorled) leaf arrangement; absence (rather than presence) of ahypanthium; subequal stamens (rather than stamens in two unequal whorls); zygomorphic (rather than actinomorphic) stamens and stigma. Under this taxonomic arrangement, Chamerion and Epilobium are monophyletic sister genera

The reddish stems of this herbaceous perennial are usually simple, erect, smooth, 0.5–2.5 m (1½–8 feet) high with scattered alternate leaves. The leaves are entire, lanceolate, and pinnately veined. A related species, dwarf fireweed (Chamerion latifolium), grows to 0.3–0.6 m tall.

The flowers have four magenta to pink petals, 2 to 3 cm in diameter. The styles have four stigmas, which occur in symmetrical terminal racemes. The floral formula is ✶/↓ K4 C4 A4+4 or 4+0 Ğ(4).

The reddish-brown linear seed capsule splits from the apex. It bears many minute brown seeds, about 300 to 400 per capsule and 80,000 per plant. The seeds have silky hairs to aid wind dispersal and are very easily spread by the wind, often becoming a weed and a dominant species on disturbed ground. Once established, the plants also spread extensively by underground roots, an individual plant eventually forming a large patch.

The leaves of fireweed are unique in that the leaf veins are circular and do not terminate on the edges of the leaf, but form circular loops and join together inside the outer leaf margins. This feature makes the plants very easy to identify in all stages of growth. When fireweed first emerges in early spring, it can closely resemble several highly toxic members of the lily family, however, it is easily identified by its unique leaf vein structure

This herb is often abundant in wet calcareous to slightly acidic soils in open fields, pastures, and particularly burned-over lands; the name Fireweed derives from the species' abundance as a coloniser on burnt sites after forest fires. Its tendency to quickly colonize open areas with little competition, such as sites of forest fires and forest clearings, makes it a clear example of a pioneer species. Plants grow and flower as long as there is open space and plenty of light. As trees and brush grow larger the plants die out, but the seeds remain viable in the soil seed bank for many years; when a new fire or other disturbance occurs that opens up the ground to light again, the seeds germinate. Some areas with heavy seed counts in the soil can, after burning, be covered with pure dense stands of this species and when in flower the landscape is turned into fields of color.

In Britain the plant was considered a rare species in the 18th century, and one confined to a few locations with damp, gravelly soils. It was misidentified as Great Hairy Willowherb in contemporary floras. The plant's rise from local rarity to widespread weed seems to have occurred at the same time as the expansion of the railway network, and the associated soil disturbance. The plant became locally known as bombweed due to its rapid colonization of bomb craters in the second world war

Because fireweed can colonize disturbed sites, even following an old oil spill, it is often used to reestablish vegetation. It grows in (and is native to) a variety of temperate to arctic ecosystems. Although it is also grown as an ornamental plant, some may find it too aggressive in that context

From Wikipedia, the free encyclopedia
Photos Troodos , 11/7/2015 by George Konstantinou